These striking images are all highly commended entries in the Wildlife Photographer of the Year competition. Now in its 54th year, the competition, run by the Natural History Museum in London, is a showcase for the world's best nature photography.
This year's competition attracted more than 45,000 entries from professionals and amateurs across 95 countries. Overall winners are announced on October 16.
American Tony Wu found this Asian sheepshead wrasse in the western Pacific Ocean near Japan's remote Sado Island. Wrasse start out as females, and when they reach a certain age and size - up to a metre long - can transform into males. This mature pastel-coloured specimen uses its pink forehead to head-butt rivals. The curious creature feeds on shellfish and crustaceans, though little more is known about it.
Isak Pretorius from South Africa found this lioness drinking in Zambia's South Luangwa National Park. Pretorius positioned his vehicle on the opposite side of the waterhole, steadying his long lens in the low light on a bean bag. The lioness appeared through the tall, rainy-season grass and hunched down to drink. With perfect timing, Isak caught her gaze and her tongue, lapping the water, framed by the wall of lush green.