Lonesome George, the last remaining Pinta Island tortoise, has died.
Lonesome George was found dead in his corral at the Tortoise Breeding and Rearing Centre in Puerto Ayora, Santa Cruz Island, by members of the Galapagos National Park Service.
Park ranger and long-time caretaker Fausto Llerena found the tortoise stretched out in the direction of his watering hole with no signs of life.
The cause of death will not be known until a necropsy has been performed. His body will be held in a cold chamber to avoid decomposition.
The age of Lonesome George is not known, but he is estimated to be more than 100 years old.