Sixth-string hooker Saia Fainga'a will start his first test match in four years as the Wallabies face a huge set-piece assignment against South Africa at Newlands.
Fainga'a becomes the fifth man to wear the jinxed No 2 jersey in eight games after gaining a shock recall ahead of Queensland teammate James Hanson.
The 27-year-old is one of three changes to the starting team which overcame Argentina 32-25 on the Gold Coast, with vice-captain Adam Ashley-Cooper returning from injury and Joe Tomane edging out Rob Horne on the other flank.
Hanson had appeared an automatic choice after the hooker's curse claimed Tatafu Polota-Nau (ankle) for the second time this year but Fainga'a's reliability in the scrums and lineouts was preferred against a Boks pack that unsettled the All Blacks in Wellington.
But the fact that Bismarck du Plessis, seen as the best No 2 in the world, is on the South African bench contrasts the rivals' fortunes.