Quade Cooper will play at first-five against the All Blacks on Saturday, with Kurtley Beale dropped from the starting XV.
However, in a departure from convention, Wallabies coach Robbie Deans has yet to announce his reserves for the test at Eden Park, a match his side must win to have any hope of taking back the Bledisloe Cup.
Cooper's inclusion after he was absent from the match-day squad in the Sydney defeat was widely predicted, as was the fact that fullback Beale faced a demotion after his mistake-ridden performance.
Brumbies flanker Michael Hooper starts in place of injured captain David Cooper, with halfback Will Genia taking over the leadership.
The return of Cooper, who suffered a serious knee injury in his last match at Eden Park - during the World Cup playoff for third and fourth - shifts Berrick Barnes out one place in the backline to second-five. Anthony Fainga'a has been dropped to make way for Barnes, who retains the goal-kicking duties.