They used to be giants, chosen for their scrum-splitting ability. The evolution of hookers, led by the All Blacks, has created a different beast - athletic ball-controllers, ball-carriers and ball-stealers - giving New Zealand a competitive edge.
Thanks to hookers like Keven Mealamu and Andrew Hore, the position has changed and arguably no one in world rugby fills the new role better than they do. Mealamu played his 101st test overnight against Italy and clocks up 10 years as an All Black this week. Andrew Hore played his 73rd test against Scotland last week - and the selection of Dane Coles can be seen as an effort to keep the hooking evolution on track.
Gone is the brutal scrum-power of hookers like England's Steve Thompson and South Africa's John Smit, the respective starting hookers of the 2003 and 2007 World Cup winners. Mealamu, Hore and now Coles clock in below 110kg and all are a mix of fourth-loose-forward mobility, ball-running, ball-snaffling and lineout accuracy. Thompson and Smit generally threatened 120kg and beyond - with 10kg making a big difference in today's high-paced, multi-phase game.
Former All Blacks selector Peter Thorburn says scrums punctuate the modern game less, meaning hookers have to be picked on a wider skill set.
"Hookers have to match flankers and the No8 by being quick away from scrums and lineouts and possessing all-round skills. That's why Coles has the edge over Hika Elliot at the moment; he is a more productive and explosive ball runner. Elliot tends to die with the ball too often and has become too readable. Smaller, mobile hookers can be handy in that regard. The Frenchman Dimitri Szarzewski plays in a similar vein; a terrific ball-carrier.