Matt LeBlanc has threatened to quit as Top Gear host if Chris Evans is not sacked.
Reports suggest that the former Friends actor is becoming increasingly frustrated with how rude his co-presenter is with the staff on set and has told producers that they can no longer work together.
The revelations are said to have rocked the BBC and could leave them having to pick between the pair, with filming for the second series due to start in September.
A show source said: "The big question now is who goes and who stays."
A BBC spokesman and a spokesman for Evans, 50, declined to comment on the reports.